Montanoa ovalifolia ssp. ovalifolia  

Montanoa ovalifolia is roadside shrub that is noteworthy for its abundant, densely clustered heads and the extreme amount of variation in a number of its characters. The leaves in particular very considerably on the individual and populational levels. In one population in Santander, Colombia (SF 5622 A-H) all of the leaf shapes for the entire species were found so there does not appear to be any correlation with locality. The other variable characters, however, do appear to have clinal variation that is here recognized on the subspecific level.

REPRESENTATIVE SPECIMENS: COLOMBIA. BOYACA: Duitama, II Dec 1969, Cuatrecasas 1693 (F,US); Oambita, 20 Oct 1972, Garcia-Barriga 20376 (COL); Chiquinquira, 9 Jul 1964, Espinal 1769 (COL); Oiiican, 18 Sep 1969, Cuatrecasas & Rodriguez 27856 (US); Tunja, 30 .Tun 1971, Guarin &Villarreal 1028 (COL); Cocuy, 14 Sep 1938, Cuatrecasas & Garcia-Barriga 1693 (COL); Tota, I Sep 1951, Yepes-Agredo 3147 (COL); Tabor, Cocuy, I Aug 1957, G/'lIbb, Curry & Fe/'l/andez-Perez 214(COL); 29 km N of Boyacii/Cundinamarca departmental limits, 15.ful 1979, SF 5579* (COL, OS); I km NE of Arcabuco, 16 Jul 1979, SF 5592* (COL, OS); 8 km S of Tunja, 20 Jul 1979, SF 5634 (COL,OS); 16 km S of Tunja, 20 .ful 1979, SF 5635 (COL, OS); 17 km S of Tunja, 20 Jul 1979, SF 5637*(COL, OS); I km S of Boyaca, 20 .ful 1979, SF 5638* (COL, OS); 5 km S of .fenesano on I'd to Tibana, 20 lui 1979, SF 5639* (COL, OS). CALDAS: 2 km N of Nerra, 30 km S of Aranzazu, 29 lui 1979, SF 5699 (COL, OS). CUNDINAMARCA: 19 km NW ofjct of I'd from Zipaquira to Urbate and I'd to Neusa,14 Jul 1979, SDF 5543* (COL, OS); 6 km N of Zipaquira on I'd to Pacho, 14 Jul 1979, SDF 5547*(COL, OS); 29 km N of Boyacii/Cundinamarca departmental limits, 15 .Tul 1979, SF 5579 (COL, OS);ca. I km SSW of Boyacii/Cundinamarca departmental limits, 15 Jul 1979, SF 5578* (COL, OS); near Sesquile, 6 Ju11947, Haught 5931 (COL, NY, US); Bogota Plateau, 27 Dec 1926, Niellleyer 152 (US); Sabana de Bogota, 20 Nov 1949, Dugand 4350 (COL, CM); Zipaquira, 18 Aug 1944, Jaralllillo s.n.(COL); Simijaca, May 1930, Perez 241 (COL); near Bogot~l., Aug 1933, Garcia-Barriga 49 (COL); Zipaquira, 20-24 Oct 1919, Pennell 2620 (OH, NY); Usme, 9 Jan 1946, Diaz 12 (US); Sabana de Bogota, .Tun 1916, Dawe 300 (K, US). SANTANDER: near Tona, 17 Feb 1927, Killip & Smith 19446 (OH, NY, US); btw California and Vetas, 15-22 Jan 1927, Killip & SlIlith 18001 (OH, NY, US); near Vetas, 16 Jan 1927, Killip & SlIlith 17885 (A, OH, NY, US). NORTE DE SANTANDER: 24 km SE of Pamplona, 4 km SE of La Legia, 18 Jul 1979, SF 5622 (OS); Pamplona, 26 Jun 1940, Cuatrecasas & Garcia-Barriga 10245 (COL, F, US).