Montanoa guatemalensis  

Trees 4-15 m tall. Stems terete, brown, herbaceous parts densely pubescent with the hairs 0.5-1.0 mm long. Leaves variable; petioles 1.0-14.5 cm long, occasionally with auricles at distal end, densely puberulent; blades ovate to pentagonal (Fig. 35A-I), 5-19 cm long, 3-18 cm wide, apex acuminate, margin entire to irregularly serrate-dentate, unlobed to 3-lobed, adaxial surface of younger leaves moderately pubescent, hairs pustular, older leaves often glabrous, abaxial surface densely glandular and pubescent, hairs whitish, giving the surface a light color, 0.5-1.0 mm long. Peduncles 1.5-2.5 mm long, densely hispid, hairs 0.5-1.0 mm long. Heads erect, 1.0-1.5 cm diam in flower, 2.0 cm diam in fruit, several to many in dense cymose synflorescences in oppositely and alternately branched compound corymbs. Phyllaries extremely variable (Fig. 4), 5-9, biseriate, subequal, reflexed in flower, ovate to ovate-lanceolate, 3-8 mm long, 2-4 mm wide, green, apex usually rounded but occasionally acute to acuminate and mucronate, margin ciliate, entire, dark, abaxial surface densely pubescent, hairs 0.5-1.0 mm long, adaxial surface glabrous. Ray.florets 6-10; corollas white, ligules obovate, 20-26 mm long, 6-9 mm wide, apex acute to 2-notched, adaxial surface glabrous, abaxial surface sparsely glandular and pubescent, hairs less than 0.5 mm long, tube 1.0-1.5 mm long, 0.5 mm wide, essentially glabrous. Disc florets (Fig. 35J) 70-100; corollas yellow, tube 0.75-1.5 mm long, 0.5-0.75 mm diam, densely glandular and pubescent, hairs less than 0.5 mm long, throat cylindrical, 2.0-2.25 mm long, 1.5-2.0 mm diam, sparsely glandular and pubescent, hairs less than 0.5 mm long, lobes 5, 0.75-1.00 mm long, 0.5 mm wide, apex acute, densely glandular and pubescent, hairs less than 0.5 mm long; stamens with filaments 2.0-2.75 mm long, 0.2 mm wide, anthers not fully exserted from corolla, thecae brown, 1.5-1.75 mm long, 0.5 mm wide, apical appendages yellow, acuminate, 0.5 mm long, 0.3 mm wide, glabrous; styles yellow, 4.5-5.25 mm long, enlarged at base, stigmatic surfaces yellow, 1.0-1.25 mm long, apical appendages yellow, long-acuminate, 0.5 mm long, 0.4 mm wide. Pales at anthesis ovate (Fig. 35K), 4.5-5.5 mm long, 2.0-2.5 mm wide, light yellow with dark bands on distal half and midrib, apex very long acuminate, dark, margin ciliate, entire, abaxial surface densely glandular and pubescent especially near the center, hairs 0.5-1.0 mm long, adaxial surface glabrous; pales at fruiting persistent, ovate to triangular (Fig. 35L), 9-11 mm long, 2.5-3.0 mm wide, indurate with prominent parallel veins, stramineous, apex narrowly tapering to a recurved, indurate tip, margin sparsely ciliate, entire, abaxial surface moderately glandular near the center and sparsely pubescent, hairs 0.5-1.0 mm long, adaxial surface glabrous; achenes brown-black, 3.5-4.0 mm long, 2.0 mm wide, smooth. Chromosome number, n = ca. 114.

Funk, V. A. "The Systematics of Montanoa (Asteraceae, Heliantheae)" in Memoirs of the New York Botanical Garden. Vol. 36. The New York Botanical Garden. Bronx, NY. 1982.