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Taxonomy
Montanoa atriplicifolia
Nomenclature
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Family: CompositaeGenus: Montanoa
SUMMARY
Verbesina atriplicifolia Pers., Syn. PI. 2: 472. 1807. Type: Hort.ltal. (holotype, P?; non-flowering isotype, FI!). Persoon often used Jussieu's herbarium (Stafleu, 1967) which is now at Paris. No possible holotypes were found in the loan material from P and Persoon's descliption is cryptic. It would not be possible to place this species except that lussieu and Desfontaines published a better descIiption (Colla, 1824), synonymy, and a drawing, and attributed the species to Persoon. The drawing is somewhat questionable, however, as the rays are too blunt for Montanoa, and they have well-developed stigmas. Although there is still some doubt, the evidence is sufficient to warrant recognition of this name. Eriocoma atriplicifolia(Pers.) Kuntze, Rev. Gen. PI. 1: 336. 1891.
Clerodendron phlomoides Juss. & Desf. in Lamarck, Hort. Ital. 8: 460. 1808, nom. nnd. pro syn.
Galinsoga discolor Spreng., Nov. Provo 19. 1819. Type: Hort. Ital., G. B. Balbis s.n. (holotype, Pi; isotype, G-DC; photo of holotype, US!; photo of G-DC isotype, GH! US!).
Montanoa dumicola Klatt in Durand & Pittier, Bull. Soc. Roy. Bot. Belgique (Brussels) 31: 200. 1882. Type: COSTA RICA, San Jose, hedgerow on the "llanos de Alajuelila," 1100 m, I Dec 1889, H. Pillier 1454 (holotype, GH! isotypes, BR[2J! G! US!).
Montanoa pauciflora Klatt, Leopoldina 23: 90. 1887. Type: "Middle America ... in Herb. Klatt." (holotype not located). The most likely place for this specimen is GH. However, the types as well as the general collections of Montanoa have been searched to no avail. Coreopsis trilobata Vahl ex Klatt, Leopoldina 23: 90. 1887, nom. nud. pro syn. This is listed as the type of M. pallciflora Klatt in the type description. The types as well as the general collections at GH have been checked for this specimen (in litt., M. Calloso) but it has not been located. The possibility exists that the specimen will be located at a later date; a neotype has not been selected.
Montanoa schotii Robins. & Greenm., Proc. Amer. Acad. Arts 34: SIS. IS99. Type: MEXICO,Yucatan, on the road between Merida and Sisal, 24 Oct IS65, A. Schall 913 (holotype, Pi)."
REPRESENTATIVE SPECIMENS: BELIZE. CAYO: EI Cayo, 12 Feb 1931, Bartlett 11421 (F, GH, MICH,MO, NA, US); Bejuco, 15 Feb 938, Gentle 2169 (F, K, MICH, US); 5 mi N of Blancaneau Lodge,12 mi S of Georgeville, 24 Jan 1974, Liesner &: Dwyer 1630 (OS); 3 mi S of Grano de Oro, 2 Jun 1973, Gentry 7761 (MO, OS); Georgeville to Augustine rd at mile 8, 3 May 1969, Proctor 30268 (BM, LL); 9 mi S of the Western hwy on rd to Augustine, 28 Jan 1970, Wilbllr &: Weaver 11491 (DUKE); Macal River 1 mi past Guacamallo Bridge, rd to Millionmio, Dwyer &: Liesner 12355 (MO); S of San Luis and E of Camp Six, 19 Mar 1967, Dwyer et al. 377A (LL, MO); Cave Branch Section, Humming Bird hwy, 18 Feb 1956, Gentle 9029 (LL). COSTA RICA. SAN JosE: San Pedro, E of San Jose, 25 Jan 1922, Green11lan &: Greell1/lall 5315 (GH, MO); San Jose, 18 Jan 1916, Holway 444 (GH). EL SAL VADOR. AHUACHAPAN: near Ahuachapan, 9-27 Jan 1922, Sa 20018 (GH, NY, US); vicinity of Ahuachapan, 16-25 Jan 1947, Standley & Padilla 2610 (F). LA LIBERTAD: stream near Colon, 21 Jan 1949, Williams & Molina 15222 (F). SANTA ANNA: just SE of pueblo of Coatepeqlle along old free rd to San Salvador from Santa Anna, 30 Nov 1978, FL 2922* (OS); 3 km SE of Santa Anna, 18 Nov 1963, Porter 1285 (DS, GH, MEXU). SONSONATE: Finca Chilata, 26-27 Dec 1921, Sa 19229 (GH, NY, US). GUATEMALA. AMITITLAN: Moran, 22 Dec 1916, Holway 625 (US). CHIMALTENANGO: above Las Calderas, 15 Dec 1938, Sa 60086 (G); Quisache, 5-6 Jan 1939, Sa 62009 (F). CHIQUIMULA: btw Chiqllimula and La Laguna, 27 Oct 1939, Se 30702 (F); hills near Ipala, 23 Oct 1939, Se 30355 (F); 2 km from village of EsquiPlilas, 6 Dec 1969, Molina & Molina 25168 (F, LL). GUATEMALA: Santa Catarina Pinula, just S of Guatemala City, 7 Dec 1977, FRa 2638 (OS); SE of Guatemala City on rd to San Salvador, 7 Dec 1977, FRa 2645* (OS). JALAPA: N of Jalapa on rd to Volcan jumay, 8 Dec 1977, FRa 2653 (OS). JUTIAPA: near EI Molina, 26 Nov 1940, Sa 78459 (Fl. PETEN: Tikal National Park, Bajo del Hormiguero, ca. 12 km on old Remate rd, 28 Jan 1961, Contreras 1887 (LL); Tikal, top of Temple IV, 5 Feb 1959, Lundell 15366 (LL); La Liberlad, 16 Mar 1935, Aguilar 476 (LL, MICH, MO, US); EI Paso, 25 Apr 1932, Lundell 1537 (MICH, US); Sacpuy, 10 km S of Flores, 28 Jan 1971, Ortiz 1555 (BM, F, MICH, US); Melchor de Mencos rd, blw kms 35-36, 5 Feb 1966, Contreras 5476 (ENCB, F, MICH, US); rd btw San Andres and Santa Elena, [0 Jan 1970, Ortiz 548(ENCB, F, MO, NY). PROGRESSO: along railroad btw Barranquillo and Cruz, 30 Jan 1942, Se 43328 (F); btw Zanarate and EI Chato bridge, 28 Nov 1969, Molina & Molina 24920 (MO, NY, US). SANTA ROSA: SE of Barberena, 7 Dec 1977, FRa 2646 (OS); SE of Barberena, 7 Dec 1977, FRa 2647 (OS); near EI Molina, 20 Dec 1936, Sa 60472 (F); E of Cui1apa, 25 Nov 1940, Sa 78257 (F); near Cuilapilla, 23 Nov 1940, Sa 78117 (F); Casillas, Nov 1892, Heyde & Lux 4242 (F, G, K, M, MO, NDG, NY). SACATEPEQUEZ: ca. 19 km N of Escuintla, 23 Jan 1977, King 7184 (MO, US); barranco above Duenas, 21 Jan 1939, Sa 63203 (F); near Tejar, 14 Dec 1938, Sa 59863 (F). HONDURAS. COMAYAGUA: near Comayagua, 12-13 Mar 1947, Siandley & Chacon 5935 (F); upstream from main vado in EI Zamorano, 2 Dec 1978, FL 2932 (OS); near Esquela Agricola Panamericana on rd to Giiinope, 4 Dec 1978, FL 2939 (OS); EI Quebracho, 29 Nov 1946, Sa 368 (F); Poza del Tacuazfn, 13 Dec 1947, Molina 671 (BH, F, GH, MO, US); near EI Zamorano, 26 Nov-9 Jan 1947, Sa 1495 (F); rd btw EI Jicarito and EI Pedregal, 13 Nov 1948, Sa 14478 (F); Rio Yeguare at Giiinope, Nov 1943, Rodriguez 1568 (F).OLANCHO: btw Catacamas and EI Hatillo along Rio de Catacamas, 27 Mar 1949, So 18866 (F); near Juticalpa, 5-16 Mar 1949, So 17409 (MICH); rd to San Francisco La Paz, 20 Nov 1963, Molilla 13309 (F, LL, NY, US). EL PARAISO: NE of Danli, 3 Dec 1978, FL 2934* (OS); ca. 1 mi N of EI Paraiso on rd to Danli, 3 Dec 1978, FL 2937 (OS). SANTA BARBARA: San Pedro Sula, Feb 1888, Thieme 5324 (F, G, GH, K, M, MO, NY, US). MEXICO: CAMPECi-IE: Campeche/Yucatan state line on hwy 261, 11 Nov 1975, Dwyer 334 (CA); 17 mi E of Xpuii1, 29 Nov 1977, FRo 2593* (OS); near Escorsega, 16 Jan 1966, Maluda 37483 (F, MEXU); km 5 on the rd from Escarcega to Candelaria, 27 Dec 1965, Herndndez el al. ES-229 (MEXU); Tuxpena, 22 Dec 1931, LUlldell 1093 (ARIZ, DS, F, GH, MICH, MO, S, UC, US). CHIAPAS: Piedra de Huixt1a, 20 Jan 1949, Maluda 18603 (F, MEXU); rd from Toliman to Niquivil near Ojo de Agua, 14 Dec 1976, Breedlove 42607 (DS); jct of Rio Perlas and Rio Jatate at San Quintin near Laguna Miramar, 15 Mar 1955, Sohns 1589 (MICH, US). QUINTANA Roo: Coba, 2 Mar 1976, Morella 535 (MEXU); Chichankanab, s.d., Gaulller 1479 (F, G, GH, K, MEXU, NY). YUCATAN: VaUadolid, 2 Jul 1932, Sleere 1688 (LL, NA, MICH); Peto, 26-27 Jul 1932, Steere 2291 (MICH); Chichen ltza, 23 Jun 1932, Steere 1488 (MICH, US); Calotmul, s.d., Gaumer 2108 (B, BM, C, F, GH, MO, NY, S); Izamal, s.d., GaulIler 2499 (F); Temax, s.d., Gaumer 2357 (BR, DS, MICH, UC, US). NICARAGUA. ESTELl: 3-7 km NW of Pueblo Nuevo, 24 Nov 1973, Williams & Molina 42393 (F). MADRIZ: mt. above Somoto, 15 Nov 1946, Williams & Molilla 10943 (F, GH,MICH, MO).
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